The parents of Ashya King, who was taken out of hospital against medical advice, face being extradited to the UK after being arrested in Spain.
Police say officers in the Malaga area pulled over the family's car at 9pm UK time on Saturday and found the five-year-old and his parents inside after a tip-off from a staff member at the hotel in which they were staying.
The boy from Portsmouth, who underwent "extensive surgery" during an operation on his brain tumour seven days ago at Southampton General Hospital, was taken to a hospital in the local area and has since been moved from a high to a lower dependency unit which Hampshire Police described as "good news".
Assistant Chief Constable Chris Shead, of Hampshire Constabulary, said his parents, Brett King, 51, and Naghemeh King, 45, were in custody after being arrested on a European arrest warrant.
In response to criticism about the force's actions, he said: "Faced with the situation that we were, we had medical experts telling us Ashya was in grave danger... if he didn't get the care that he needed there was a potential threat to his life.
An international search was launched after Ashya was taken on Thursday "So I make no apologies for being as proactive in this investigation as we have been.
"There are no winners in this situation. I've said all along that this must be a terribly distressing time for Ashya's family and I stand by that now."
He added that it was too soon to say when Ashya would come back to the UK, but said Southampton General Hospital had been contacted and was liaising with doctors taking care of him in Spain.
Ashya's brothers and sisters were not in the vehicle, and found in a hotel about 10 miles away.
Mr Shead said a team of Hampshire police officers would be arriving in Spain later to continue the investigation, as proceedings to extradite Mr and Mrs King to the UK get under way.
French media had picked up the story after Ashya was taken from hospital The arrests came as footage emerged on video-sharing website YouTube in which Ashya's father, a Jehovah's Witness, insisted they had taken him from hospital to seek a cancer treatment not available on the NHS.
"We were much disturbed today to find that his face is all over the internet and newspapers and we've been labelled as kidnappers, putting his life at risk, neglect," he said.
"As you can see, there's nothing wrong with him. He's very happy actually, since we took him out of hospital. He's been smiling a lot more, he's been very much interacting with us.
"But I just wanted to say very quickly why we took him out of the hospital.
Ashya is now being cared for in a Spanish hospital "The surgeon did a wonderful job on his head that took out the brain tumour, completely they reckon. But straight away afterwards he went into what's called posterior fossa syndrome, which means very limited moving or talking or doing anything."
He said he had spoken to specialists after Ashya's surgery and had requested proton beam treatment, which was not available on the NHS.
"Proton beam is so much better for children with brain cancer," he said.
"We pleaded with them for proton beam treatment. They looked at me straight in the face and said with his cancer - which is called medulloblastoma - it would have no benefit whatsoever.
"I went straight back to my room and looked it up and the American sites and French sites and Switzerland sites where they have proton beam said the opposite, it would be very beneficial for him."
Jehovah's Witnesses refuse blood transfusions on religious grounds but are open to other medical procedures.
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